Tag Archives: hemp

Studies in laboratory animals have shown CBD to protect the liver from toxic insults

Assay of the extract was 61% edible fatty acids, 26% phytocannabinoids and 13% other plant chemicals including fatty alkanes, plant sterols, triterpenes, and tocopherols. In the 14-day repeated oral dose-range finding study reported by Marx et al., a No Observed Adverse Effect Level could not be determined, however, the results of a 90-day repeated dose study with a 28-day recovery period in Wistar rats was also reported. In this study, doses of 0 , 100, 360 and 720 mg extract/kg bw per day were used. Significant decreases in body weight, body weight gain, and differences in various organ weights, compared to controls, were reported at the mid and high dose levels, but the authors concluded that many of the findings were reversible as they were trending towards normal at the end of the recovery period. A NOAEL for the hemp extract in Wistar rats in the 90-day study was determined to be 100 mg/ kg bw per day and 360 mg/kg bw per day for males and females, respectively. In the 90-day study being reported here, test article related significant changes in body weights, daily body weight gain and feed efficiency were seen in the males in all treatment groups which was still noted at the end of the recovery period. The magnitude of the significant change in body weights, daily body weight gain and feed efficiency in the low and mid dose groups was less than 10% and showed signs of obvious recovery and were therefore considered to be not toxicologically relevant. The effect in the males receiving 800 mg/kg/day was >10% and was still evident at the end of the recovery period and was considered toxicologically relevant. Reported rodent studies have differing findings on hepatotoxicity when CBD is orally administered in high doses. Hepatocellular hypertrophy with a centrilobular pattern was observed in rat livers in the study being reported.

This pattern of hepatocellular hyperplasia is frequently observed in rats and other animals exposed to agents that induce the CYP family of enzymes and can be associated with activation of peroxisome proliferator-activated receptors. THC has affinity for PPARα,trimming tray and CBD has very low to no affinity for PPARα and high affinity for PPARγ. Interaction with the PPARγ is one of the mechanisms of action for CBD. In our study, we did not show the mechanism of action for the hepatocellular hypertrophy. We did show that the activities of liver enzymes in serum were not significantly changed by treatment with the test article and the hepatocellular hypertrophy was reversed during the 28-day recovery period. In the study reported by Marx et al., no histopathological changes were observed in the livers from the treated and control rats and the liver weights in the male and female rats in the 360 and 720 mg/kg body weight/day were significantly increased at 90 days. The 28-day recovery males and females receiving 720 mg/kg/day retained the significantly increased in hepatic weights. The induction of hepatic drug metabolizing enzymes can be associated with increased liver weights, and hepatocellular hypertrophy and hyperplasia and elevation of hepatic-source enzymes in serum. The evidence in the scientific literature supports a conclusion that the centrilobular pattern of hepatocellular hypertrophy and increased liver weights observed in our study was due to induction of HDMEs and/or peroxisomes. No hepatocellular necrosis and changes in the clinical chemistries occurred which is evidence that liver damage did not occur. This conclusion is further supported by not observing hepatocellular hypertrophy and increased liver weights in the 28-day recovery groups that received the test article.In the study being reported both the treated and control male rats had the same incidence and severity of vacuolization of the adrenal zona fasciculata and the adrenal weights were significantly increased in the Group 4 females. The vacuolization of the adrenal zona fasciculata and increased adrenal weights were not observed in Groups 5 to 8. The histopathological lesions noted in the adrenal glands in the current study was seen in both control and high dose males and is not considered to be due to treatment with test article and not toxicologically relevant. The hemp extract in these studies was shown to be non-mutagenic in a bacterial test system used to evaluate mutagenicity. Marx et al. reported on a GLP-compliant study that concentrations of 5, 000 μg/plate of a CO2 supercritical extract of C. sativa were not mutagenic in a bacterial test system. Our GLP-compliant mutagenicity testing on the diluted extract showed that concentrations of 76,355 μg/plate were not mutagenic with and without the S9 metabolic activation. The extracts produced by isopropanol extraction and supercritical CO2 extraction were not mutagenic with and without S9 metabolic activation at concentrations up to 5000 μg/plate. The bacterial test system with the S9 mix did cause mutagenicity providing evidence that mutagenic metabolites were not produced with any of the extracts. The two additional Ames tests conducted on the undiluted extracts produced by two different extraction methods, were conducted to determine if the method of production or the olive oil diluent impacted the results of the Ames assay. No mutagenicity was noted in any of the tests conducted. Other botanical extracts have been evaluated for mutagenicity. Mutagenic studies on extracts from the plant Euphorbia triaculeata showed that it is not mutagenic and provides protection from the mutagenic effects of cyclophosphamide.A study on a novel taste modulating powder derived from Cordyceps sinensis showed this product was not mutagenic in the Ames test and these results were supported in the micronucleus assay. In a study on the genotoxicity of CBD in Caco-2 cells, 10 μM of CBD did not significantly cause DNA damage after 24 hours of incubation, and CBD was also shown in the comet assay to protect Caco-2 cells from hydrogen peroxide-induced DNA damage. CBD at an oral dose of 1 mg/kg was shown to significantly reduce azoxymethane-induced colonic aberrant crypt foci, colonic polyps and tumors. In summary, the test article, both undiluted and diluted in olive oil, was not mutagenic in a bacterial reverse mutation assay and the NOAEL in the 90-day study was concluded to be 800 mg/kg bw/day and 400 mg/kg bw/day for female and male Sprague Dawley rats, respectively. This assessment adds significant data to the currently available literature as to the safety and toxicology of CBD rich hemp extracts. Given the potential of CBD for a variety of human uses and the limited data currently available, these results support that hemp extracts are likely safe human consumption and additional studies should be conducted to validate this conclusion. Natural lignocellulosic fibers are in past decades gaining increased attention as sustainable materials for polymer composite reinforcement in substitution of energy intensive and non-recyclable synthetic fibers. Indeed, NLF reinforced composites are being considered for applications in civil construction, food packing, automotive components  and ballistic armor. It is worth mentioning that nanocellulose fibrils obtained from NLFs are being investigated as possible reinforcement for novel bio-nanocomposites with special properties for medical applications as well as production of biodegradable plastic films. A well known NLF, the hemp fiber, has been for decades extensively reported as reinforcement of polymer composites in numerous articles and mentioned in most review papers. In fact, hemp fiber/polypropylene has been used as automotive parts. The remarkable mechanical properties of the hemp fiber, tensile strength of 900 MPa and modulus of 70 GPa, offer a possibility of use in ballistic armor reinforcing stronger thermoset polymeric matrices such as epoxy and polyester. This ballistic application has not yet been investigated in hemp fiber composites. Therefore, the objective of the present work is, for the first time, to compare the mechanical properties, evaluated by bend and tensile tests, of both epoxy and polyester composites reinforced with different amounts of hemp fibers. This would allow a definition, in terms of matrix and incorporated volume fraction, of the most suitable composite for application in multilayered armor system.The hemp fibers, Cannabis sativa, investigated in this work were supplied by Designan Fibras Naturais, Brazil, as a bundle illustrated in Fig. 1. Fibers were separated, washed in running water and dried at room temperature for one week. Characterization of the hemp fibers was conducted prior to their processing in composites. One hundred fibers were randomly pick up and measured in a profile projector Nikkon, Japan, to determine the equivalent mean diameter. The average density was obtained by the mass, in a 0.001 g precision scale, divided by the volume considering cylindrical fiber. Both epoxy and polyester resins were purchased from Resinpoxy, Brazil. The epoxy was a diglycidyl ether of the bisphenol A hardened with stoichiometric phr 13 of triethylene tetramine as the catalyst. The polyester was an unsaturated ortoftalic resin hardened with 2 wt% of methyl ethyl ketone for the curing process.Fabrication of the composites for mechanical tests was carried out by laying up continuous and aligned hemp fibers inside a steel mold, Fig. 2, trim tray pollen with internal volume of 150 × 120 × 7mm. Fibers were first cut and weighed according to the volume fraction, calculated by considering their previously evaluated density as well as the densities of epoxy and polyester from. Just before processing, the fibers were dried in a stove at 60 ◦C for 3h. Layers of fibers were placed inside the mold together with either epoxy or polyester resin, already mixed with corresponding hardener. After closing the mold, a pressure of 1 ton was applied and left to cure for 24h at RT. The produced composite plate, Fig. 2, was machined into 150 × 10 × 7mm prismatic specimens for 3 points bend tests as per ASTM D790 standard. Flat tensile specimens were directly fabricated in a specially designed steel mold, Fig. 3, in which hemp fibers were aligned together with the chosen resin already mixed with corresponding hardener. Shape of specimens with 7 × 7mm of gage cross section and 20mm of gage length as per ASTM D 638 standard. A pressure of approximately 1 MPa was applied to the mold’s lid during the cure at RT for 24h. Both flexural and tensile tests  were carried out in a model 5582 Instron machine, USA, operating at RT and a crosshead speed of 0.5mm/min. Fracture analysis of ruptured specimens was performed by scanning electron microscopy in a model SSX-550 Shimadzu microscope, Japan, operating with secondary electron at 20 kV. Fourier transform infrared analysis was performed to detect functional groups in the hemp fibers as well as the effect caused by addition of these fibers in the epoxy functional group. Band spectra were obtained in a model IR-Prestige- 21 Shimadzu spectrometer, Japan, using 2mg pressed ground samples mixed with 110mg of KBr.Frequency histogram for the equivalent diameter of the as-received hemp fibers is shown in Fig. 4. The average length is 76.6mm, while the average diameter, Fig. 4, was found as 65 m. The density of the fibers was obtained as 1.35 ± 0.27 g/cm3 with thinner fibers presenting comparatively higher densities. Fig. 5 shows FTIR spectra of the hemp fiber and three hemp fiber/epoxy composites. The hemp fiber spectrum in Fig. 5 displays intense broad bands centered at 3333 and 2916 cm−1 that could be associated with O H and C–H groups in the fiber cellulose. The sharp band at 1738 cm−1 might be assigned to the C O stretching of carboxylic group of hemicellulose, while the band at 1656 cm−1 was indicated as C C as stretching of unsaturated acids or sterols in tannin. The band at 1510 cm−1 was associated with the aromatic skeletal vibration of lignin. Bands at 1425 and 1377 cm−1 and around were attributed to C–H bending as well as C–H2 rocking vibration of Fig. 4 – Histogram for equivalent diameter of the as-received hemp fibers. groups in lignin and cellulose. The 1160 and 1030 cm−1 bands were assigned to cellulose, respectively, C–O C asymmetric valence vibration and C–O stretching primary alcohol. Finally, the band at 897 cm−1 corresponds to C–H rocking vibrations of cellulose. The FTIR spectra of the hemp fibers/epoxy composites in Fig. 5 revealed bands corresponding to the plain DGEBA/TETA epoxy such as: 2963 and 2866 cm−1 assigned to C–H alkyl group; 1612 and 1510 cm−1 assigned to C C phenyl ring; 1456 and 1257 cm−1 assigned to H C–H groups; 1255 and 1030 cm−1 assigned to C–O C ether; and a small 915 cm−1 band assigned to vibration of epoxide group.

Stem elongation and fiber development are associated with elevated levels of gibberellins

Without shattering,immature seeds would all fully mature,increasing yield by up to 15%.Thus,further domestication of non-shattering cultivars could greatly improve yield via a multifold mechanism,preventing harvesting loss and permitting all seeds to reach maturity.Seed traits that expand market options will also be valuable.For example,there has been little research investigating the differences in hemp seed flavors.Taste tests in our lab identified varieties with weak to strong flavors of hazelnut or walnut ,as well as one with a mild flavor.More work has been done on altering seed oil composition,although hemp seeds already possess valuable v-3 characteristics.Hemp seed oil is 85% polyunsaturated fatty acids with 60% and 24% being v-6 and v-3 fats,respectively.Further increases in v-3 fatty acid levels might foster the favorability of hemp seed for human and animal dietary needs.Overall,different tastes and oil compositions would expand the use of hemp seed in human and animal food products.Hemp stalks contain two key fractions,the bast fiber and hurd.To separate bast fibers from the inner hurd,the stalks must undergo a process called ‘retting’.Retting relies on the diverse microbial populations in the environment to break down pectin and other components that bind the fibers to the hurd tissue.Crop maturity at harvest,retting method,environmental conditions,as well as the nature of the bacterial and fungal populations,are factors that impact retting.Harvesting the crop at initiation of flowering improves fiber yield,strength,and quality.Continuing studies on the biodiversity,relationships,and functions of microbial communities will improve our understanding of the retting process and augment the consistency of obtaining high quality products.Retting methods,ebb and flow primarily dew- and water-retting,pose drawbacks,including inconsistent fiber strength and quality,and polluted wastewater,respectively.

Development of varieties having bast fiber with higher cellulose content as well as lower pectin and lignin cross-linkages may decrease the retting requirements,thus improving fiber strength and quality while saving time and labor.Hemp produces a diverse array of nonintoxicating phytocannabinoids,terpenes,and phenolic compounds with potential pharmaceutical values as drugs or supplements.The biosynthesis of terpenophenolic phytocannabinoids in Cannabis is well understood,albeit,several early steps in the pathway remain to be characterized.Understanding the regulation of phytocannabinoid biosynthesis is vital to development of varieties that are optimized for production of desirable metabolites while maintaining low levels of THC.Little is known about the endogenous and environmental regulation of phytocannabinoids.Abscisic acid,ethylene,and gibberellic acid modulate the production of phytocannabinoids.However;at present,factors controlling the epigenetic,transcriptional,and post-transcriptional regulation of phytocannabinoid biosynthesis remain uncharacterized.Hemp trichomes are classified into bulbous,capitate-sessile,capitate-stalked,and nonglandular types.Phytocannabinoid production and accumulation are localized to the capitate-stalked glandular trichomes.Increased production of phytocannabinoids in marijuana is,at least partially,due to the presence of larger glandular trichomes.Elucidating hormonal and other signaling cascades that regulate the development and size of specific types of hemp trichomes will also be important in maximizing phytocannabinoid production in hemp.The effects of agronomic practices and nutrients on phytocannabinoid production also need to be investigated.Anecdotal claims from marijuana growers suggest that pollination of Cannabis flowers lowers phytocannabinoid yield,consistent with decreases in essential oil levels.Further studies to evaluate this concern are essential to maximize the production of CBD and other desired phytocannabinoids.Germplasm collections are a fundamental source of genetic and phenotypic diversity for plant breeding and research.Currently,access to and utility of accession collections remain limited due to the lack of a core Cannabis germplasm collection.

As THC levels may limit germplasm utility in many regions,accessions with <0.3% THC should be used to form a hemp-only germplasm core collection.Establishment of a core collection encompassing the range of hemp genetic and phenotypic diversity would increase the utility of germplasm resources and be invaluable for breeding and genetic analyses.Comparisons of accessions present in existing collections are needed to help establish such a core collection.Similarly,centralized and curated collections of hemp mutants are not available.The development of mutant germplasm collections will provide a rich source of genetic variation for studying gene function and improved traits for breeding.Hemp is a dioecious plant with female and male hemp plants being valued differently depending upon the products.For phytocannabinoid production,a pure female population is most desirable.As a seed crop,a female predominant population,with a limited number of male plants for pollination,or a monoecious variety,is most desirable to maximize yield.For fiber production,males and females are both utilized,although males are preferred.Therefore,a major goal of hemp growers and breeders is to quickly and easily determine or manipulate the sex of plants,preferably prior to planting.Sex in hemp is genetically determined by a pair of heteromorphic sex chromosomes; females have an XX chromosome pair whereas males have XY.However,environmental conditions and phytohormones can affect sexual phenotype,suggesting other overriding regulatory mechanisms are involved in determining sex in hemp.Monoecious cultivars possess XX sex chromosomes,but they produce flower clusters with male flowers at the bottom and females towards the top of each inflorescence.Notably,male flowers occur as the plant transitions from rapid growth to flowering.In hemp,gibberellins are associated with plant masculinity and greater fiber number,length,and diameter.Thus,a concentration gradient of gibberellin and other hormones may dictate inflorescence sex.Genetic markers have been developed to differentiate sex in hemp plants; however,such a method is not viable for commercial plantings.Recently,quantitative trait loci were identified for sex expression in dioecious and monoecious hemp.

Cloning of the responsible genes from these QTLs will greatly improve our understanding of genetic control of sex in hemp.Identification of genes present on the sex chromosomes,especially outside the pseudoautosomal recombinant region,will be critical for understanding sex-linked traits.Continued development of molecular markers is needed to improve QTL mapping resolution and for marker-assisted selection of desirable traits in breeding programs.The organic food market is a key player in promoting hemp food and CBD products.As such,widespread public acceptance of transgenic hemp is unlikely.It also remains unknown whether the public will welcome hemp modified using gene-editing techniques,which lack non-plant or plant-pest DNA sequences.Thus,many improvements to hemp will probably be accomplished using traditional breeding methods.However; for research purposes,the development of applicable molecular biology techniques is imperative to further study the molecular mechanisms that determine important traits in hemp.Publication of a draft-quality Cannabis genome and other genetic studies have shed some light into the difference between marijuana and hemp.The Cannabis draft genomeAppendix Aiii has been compared with draft genome sequences of its closest relative common hop as well as more distant species including bread nut and mulberry.Recently,low coverage whole-genome sequencing and genotyping-by-sequencing have been performed on 54 and 325 distinct cultivars,respectively.However; with only raw data files available,the lack of websites with easy to-use graphical user interfaces for data analyses limits the utility of these draft-level genome sequences.Transcriptome assemblies are also available ,but are primarily targeted toward understanding phytocannabinoid metabolism.Comparison of marijuana and hemp indicates that the expression of phytocannabinoid biosynthetic genes is higher in marijuana,suggesting that transcriptional regulation of the pathway may be one factor controlling cannabinoid production.Recently,a transcriptome was generated for hemp bast fibers at different growth stages,providing insight into fiber development.The evolution of genetic differences between seed/oil,fiber,and dual-purpose cultivars is less studied.In-depth genetic comparisons of diverse seed/oil,fiber,and phytocannabinoid cultivars are needed to identify the specific genes and mechanisms controlling important yield traits.

To attain the full benefit of these and other studies across species,the genome sequence needs to be improved beyond draft quality,and websites with user-friendly graphical user interfaces must be developed.To characterize hemp gene functions,methods to manipulate gene expression are urgently needed.Protocols for developing transformed hairy roots and cell suspension cultures are available,but the utility of both methods is limited since neither tissue produces seed,fiber,or phytocannabinoids.A whole-plant regeneration protocol has been developed for marijuana,suggesting that the development of transgenic hemp plants is feasible.Virus-induced gene silencing methods would also prove useful for studying gene function,but thus far have been unsuccessful.Alternatively,isolation of mutants from chemical mutagenesis screens is possible,but extremely difficult due to the anemophilous and dioecious nature of hemp.Currently,exploitation of the natural genetic diversity present within hemp may be the most straightforward way to study gene functions.In the present study,salivary biomarkers for gastric ulceration in pigs were explored,and significant differences in oxylipin levels were observed between healthy pigs and pigs with gastric ulcers.The results suggest that reduced levels of oxylipins deriving from linoleic acid might function as biomarkers for gastric ulcers in pigs.The potential of oxylipins as biomarkers for early diagnosis of gastric ulceration and thereby as a tool to prevent severe gastric ulcer development would be relevant investigating.Moreover,hemp seed hulls might reduce the incidence of severe gastric ulcers in pigs.Ulceration of the stomach is a common disease in pig production worldwide and is associated with economic losses as well as animal health and welfare issues.Severe ulcers often lead to weight loss,and bleeding ulcers may lead to spontaneous death.Thus,prevention of gastric ulcers is important to improve animal production as well as animal health and welfare.The structure of the feed is known as a major risk factor for the development of gastric ulcers.Several studies have observed an association between fine-pelleted feed and incidences of gastric ulcers,presumably due to a more fluid stomach content,dry racks which increases the risk of the acidic chyme of the more distal regions coming in contact with the non-glandular part of the stomach.Thus,feeding ingredients with a high swelling capacity is expected to result in a more solid stomach content and thereby reduce the risk of developing gastric ulcers; however,there is a gap in knowledge regarding potential dietary treatments.Hemp is an old culture plant which is easy to cultivate and manage and is known as an important source of fibre as well as a medical product due to its high content of bio-active compounds with antioxidant properties.Different hemp feed products exist,including hemp seed cake and hemp seed hulls.Hemp seed cake is a by-product after oil extraction and has a high protein content as well as some fibre,whereas hemp seed hulls are the outer shell of the seed with a high fibre content.Studies indicate that hemp might protect the gastric mucosa by inhibition of gastric acid secretion through cannabinoid receptors in the gastrointestinal tract including the stomach.In addition,the potential swelling capacity of hemp seed hulls may result in a more solid stomach content.

Clinical diagnostics of gastric ulceration in pigs is difficult since gastric ulcers are often sub-clinical with only the most severe ulcers resulting in the development of clinical signs,such as anorexia,anaemia or sudden death.Therefore,an efficient non-invasive method is required to be able to detect early stages of gastric ulceration in pigs,allowing intervention with preventive measures,e.g.by regulating the particle size of the feed,to reduce the economic losses as well as animal health and welfare issues.Saliva is a bio-specimen that has received attention for detection of disease biomarkers,as it has shown to contain compounds that are associated with certain diseases.In addition,collection of saliva is simple and noninvasive.Salivary glands with high permeability are surrounded by capillaries allowing exchange of molecules.Hence,saliva contains a variety of locally synthesised compounds as well as systemic compounds that might act as biomarkers.Studies have indicated that several factors present in saliva protect the oesophagus,including epithelial growth factors,mucins and prostaglandins.The maintenance of oesophageal health is to a large degree dependent on saliva ,indicating that saliva might be a potential biospecimen for identifying possible biomarkers that reflect the health status of the upper part of the gastrointestinal tract,including pars oesophagea.Since gastric ulcers in pigs are mostly located in the region of pars oesophagea,saliva might contain metabolites that indicate the presence of gastric ulcers.Non-targeted metabolomics is an analytical technique used to study a wide range of small molecules in biospecimens,making it possible to discover metabolites that discriminate across phenotypes,including health status.In this study,the saliva metabolome was explored for potential biomarkers associated with the presence of gastric ulceration in pigs.In addition,the effect of hemp on the incidence of gastric ulcers was examined.It is hypothesised that saliva contains metabolites that can be used as biomarkers for gastric ulceration.Furthermore,it is hypothesised that hemp seed cake and hemp seed hulls reduce the incidence of gastric ulcers via anti-inflammatory effects and by conferring more solidness to the gastric content,respectively.The pigs were provided dry feed ad libitum in single-space feeders and were offered the experimental diets from approximately 30 kg BW and until slaughter weight.One dry-feed dispenser with water supply and one drinking trough were available per pen.All pigs had permanent free access to water.Four experimental diets were formulated: meal feed without addition of hemp seed products ; pelleted feed without addition of hemp seed products ; pelleted feed added 4% hemp seed cake ; pelleted feed added 4% hemp seed hulls.Ingredient composition of the experimental diets is shown in Table 1.